Female Euodynerus quadrifasciatus, one of our rarest solitary wasps found chiefly in Devon and Dorset

Glanville Fritillaries (Melitaea cinxia) mating The wasp Ancistrocerus scoticus preys on Lepidoptera or beetle larvae Male Oxybelus argentatus, a nationally scarce wasp
Female mining bee Dasypoda hirtipes showing the 'hairy' legs for which they are famous Nests are dug by swiftly 'paddling' the earth backwards After a lengthy absence Andrena vaga is nesting in southern England again
Andrena vaga female returning with abundant willow pollen to her nest at Dungeness RSPB reserve Female Megachile circumcincta, which is now concentrated on the coast Male leaf-cutter bee Megachile maritima
Female cuckoo bee Coelioxys conoidea, which preys on Megachile maritima Bombus humilis worker Bombus sylvarum (Shrill Carder Bee) worker
Male Bombus muscorum Queen Bombus ruderarius Male Bombus ruderatus
Red Clover is an ideal foraging plant for the Short-haired Bumblebee queens released at Dungeness Part of the area nurtured for the Short-haired Bumblebee at Dungeness The Dune Robber-fly (Philnocus albiceps)
Glow Worms (Lampyris noctiluca) benefit from unimproved grassland Glow Worm eating a small snail Dune Chafers (Anomala dubia) mating
Black Marram Weevil (Otiorhynchus atroapterus) The Dune Tiger Beetle (Cicindela maritima) is nationally scarce The Cliff Tiger Beetle (Cicindela germanica) is a BAP and Red Data Book 2 species
11-spot Ladybird (Coccinella undecimpunctata) Bibio pomonae is uncommon in southern England Bibio leucopterus flies in the spring
The hoverfly Helophilus trivittatus is not that common but turns up in coastal areas as much as any habitat Six-spot Burnet moth (Zygaena filipendulae) on Sea Holly The Ground Lackey (Malocosoma castrensis) is now found almost exclusively in eastern coastal counties

Certain species of invertebrate are specific to the coast, others are also found inland but find sand dunes in particular suit their requirements. More are shown in the Purbeck pages in this section, but here are a few photographed elsewhere.

Glanville Fritillary butterflies (Melitaea cinxia, wingspan 45mm) are a Biodiversity Action Plan priority species, principally because their breeding requirements are so precise, leading to their being restricted to the Isle of Wight.

Essentially they need unstable cliffs or coastal slopes where the caterpillars' foodplant, Ribwort Plantain (Plantago lanceolata), can thrive. The Isle of Wight offers this in abundance, and in June there is usually a plentiful supply of adults. However, given the necessarily temporary nature of any breeding habitat based on erosion, there may be problems for the Glanville Fritillary in the long-term.

Euodynerus quadrifasciatus (13mm) is a very localised solitary wasp found only in southern England and mainly on the coast at Sidmouth in Devon and on the Isle of Portland in Dorset. It seems to nest in rocks and pebbles in these locations. Elsewhere in Europe, and probably in Surrey where there are records from 2002, it mostly uses wood. Prey consists of micro-moth caterpillars and/or leaf beetle larvae.

Like Euodynerus quadrifasciatus Ancistrocerus scoticus (12mm long) is one of the potter/mason wasps and is foundPellenes tripunctatus male, a Red Data Book 1 species mainly on the coast. They prey on small Lepidoptera caterpillars or beetle larvae to stock the nest, which is usually constructed in hollow places such as crevices in rocks or in dead plant stems.

The solitary wasp Oxybelus argentatus (6mm) is a nationally scarce sand specialist. They are found in south-east England in a few pockets on the coast and in Norfolk, as well as in a handful of inland heaths. Their behaviour is similar to Oxybelus uniglumis, dealt with under Sandpits in the Lowland Heathland section of this website.

In July and August coastal dunes are a classic setting for one of the most attractive of the mining bees, the nationally scarce Dasypoda hirtipes (12mm), though they are also found in inland heaths.

The female has the largest scopae (pollen brushes) of any British bee and can carry formidable quantities of pollen collected from Composites. Using the hind legs almost like paddles to remove the spoil, she swiftly digs a burrow up to 60cm in depth. This contains several cells, each stocked with around 300mg of pollen rolled into a ball and supported on three plinths of sand to allow air to circulate and maintain quality while the larva in the cell develops. The larvae pupate the following year.

Equally large, and equally impressive, is the mining bee Andrena vaga (12mm), which has returned to Britain after an absence of more than 60 years. It takes pollen only from willow trees, and initially was seen though not identified at Dungeness RSPB reserve in 2009. Since then it has significantly increased the colony in a sandy bank at Dungeness, has been found in Hampshire and has spread into inland Kent. This is a large solitary bee so further expansion is hoped for.

Among leaf-cutter Megachilid bees, Megachile leachella (12mm), formerly Megachile dorsalis, is dealt with in the Purbeck pages of this website, but another member of the genus found mainly though not exclusively on the coast is Megachile maritima (14mm), which is locally common. This species is preyed on by the physically impressive cuckoo bee Coelioxys conoidea (13mm). From the same genus, Megachile circumcincta is becoming scarce and local. They used to be found inland but now are pretty well restricted to coastal dunes.

On a slightly larger scale, the coast, particularly Kent and Essex, now provides one of the last strongholds for several species of bumblebee. Considering their status as one of Britain's most obvious, best-known and best-loved invertebrates, the bumblebee family is in a pretty parlous state, to some extent due to modern agricultural practices resulting in the removal of flower-rich grasslands. All but one of the species shown here, three of which are the subject of Biodiversity Action Plans, were seen in Kent on one day in July 2008.

Bombus humilis (14mm) has small nests with fewer than 100 workers and is associated with grasslands holding legumes, especially clover. Bombus sylvarum (12mm), or the Shrill Carder Bee, is a small species and one under severe pressure,The Bridge Spider (Larinioides sclopetarius) sets up home in the hides at Dungeness being found in only five areas of the UK. They, too, use legumes and also labiates. Bombus muscorum (15mm) needs similar flowers and although more widespread is also declining. The same comment applies to Bombus ruderarius, (16mm), which closely resembles the Red-tailed Bumblebee. They use old mouse or vole burrows for their nests. Bombus ruderatus (18mm) is a large species found only in south-east England, and not often there. It may be associated with wetland areas and nests are quite large, reportedly holding at least 150 workers. More research about, and assistance for, all these species is certainly required.

Bombus subterraneus, the Short-haired Bumblebee (15mm), was not photographed along with these other bumblebees in 2008 for a good reason. The species had been declared extinct in Britain 20 years earlier but, backed by various conservation groups, the campaign to try and reintroduce the species resulted in dozens of queens being taken in Sweden and released at the Dungeness RSPB reserve in Kent in May 2012 then again in June 2013. This was the last place where the bee had been seen in the 1980s. It took a huge amount of effort to get the habitat in prime condition, full of appropriate flowers such as Red Clover, for the release. One can only hope that the reintroduction proves successful but the bad summer of 2012 did not help in that regard. Equally, all the excellent foraging areas in Kent that have been developed over the last few years to help this bumblebee will undoubtedly assist other bees too.

Glow Worms (Lampyris noctiluca) are fairly large beetles which have suffered from the so-called improvement of grassland in the modern era and are at least as common inland as on the coast. However, my first sighting of any for many years was at Sandwich in Kent in 2008, hence their inclusion on this page. The winged male is 10-12mm and the wingless female 15-20mm. They are active mostly after dark and so need good cover to conceal themselves from predators. Grassland vegetation, logs, cracks and crevices with humid conditions are ideal habitats. In addition, the female needs a prominent position such as a bank, hummock, tussock or similar feature, to attract a passing male.

The green light females use is produced by a series of chemical reactions within the cells of the light organ. Oxidation of an active chemical, luciferin, in the presence of a catalyst, luciferase, causes the shedding of atoms and a release of energy in the form of light. The reaction is remarkably efficient since only 2%The Dune Wolf Spider is locally common but appears to have declined of the energy is lost as heat. As a consequence a brilliantly glowing female remains cold to the touch. Adult Glow Worms are active between June and August in a short summer breeding period. Snails are among the food taken.

Among other invertebrates, the Dune Tiger Beetle (Cicindela maritima, 13mm) looks very like the Heath Tiger Beetle (Cicindela sylvatica) shown in the Heathland pages of this website but as the name suggests, it is a coastal species occurring on dunes and sandy beaches. Within the area covered by this website that is in Kent and Norfolk, though Hampshire also used to host the species. Classified as Nationally Scarce, like most of the others in the genus it runs and flies readily in search of invertebrate prey. Breeding occurs in spring and summer, with larvae in burrows in hard-packed sand.

Even rarer is the smallest of the genus, the Cliff Tiger Beetle (Cicindela germanica, 7mm), a BAP and Red Data Book 2 species. This species does not fly habitually and hunts small invertebrates such as ants. Formerly widespread along the coast between Hampshire and Devon, and also present in South Wales, since 1970 it has been recorded only in Dorset and on the Isle of Wight.

The Dune Robber-fly (Philnocus albiceps, 12mm) lives up to its name by foraging for varied prey around sand dunes, using the proboscis to suck victims dry. Other invertebrates which are particularly at home in dunes are the Black Marram Weevil (Otiorhynchus atroapterus, 6mm) and the Dune Chafer (Anomala dubia, 9mm). The 11-spot Ladybird (Coccinella undecimpunctata, 6mm) is not so particular, but coastal areas are among the best places to look for it. This also applies to the flies Bibio lecopterus (12mm) and, in the south at least, Bibio pomonae (12mm). The latter, distinguished by the brilliant colouring on the femora, is found much more easily in northern England.

The handsome hoverfly Helophilus trivittatus (16mm) is the largest of the five-strong genus in Britain and though not common it is widely distributed, with coastal areas leading the way. The species visits flowers to take nectar, including thistles and umbellifers such as Hemlock Water-dropwort as in the image. The resident population is probably increased by migrants each summer.

The Six-spot Burnet moth (Zygaena filipendulae, wingspan 45mm), the commonest of seven Burnet moths in Britain, is often found on sandhills. Much rarer, and now virtually limited to eastern coastal counties, is the Ground Lackey moth (Malocosoma castrensis, wingspan 35mm). The larvae feed on saltmarsh plants including sea-lavender.

Among spiders, the undoubted star is the small but very smart crab spider Pellenes tripunctatus (6mm), which is found only on coastal shingle at Dungeness, Rye Harbour and Chesil Beach. This is a Red Data Book 1 species, although since its sites are all protected it can hardly be regarded as in danger of extinction unless permanently rising sea levels have an impact.

Dungeness is also home to the Bridge Spider (Larinioides sclopetarius, 12mm), which is widespread in the area covered by this website and is associated with solid structures close to water, including bridges, bird hides, fences and tree trunks. In contrast, the Dune Wolf Spider (Xerolycosa miniata, 7mm) can be seen scurrying across the sand from May to August.

Images © Jeremy Early. All rights reserved.

In 2013 I published My Side of the Fence - the Natural History of a Surrey Garden. Details may be found, and orders placed, via this hyperlink My Side of the Fence. In November 2015 Surrey Wildlife Trust published the atlas Soldierflies, their allies and Conopidae of Surrey, jointly written by David Baldock and me. Details are on this web page: Atlas.